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Donkey Kong and his friends decided to take a well-earned vacation on beautiful Sun Sun Beach, located, of course, on tropical Sun Sun Island. After enjoying a splash in the ocean, a hungry DK and his friends saw a massive banana floating atop a mountain. Without a moment\'s hesitation, DK up and raced for the mountaintop. Who knows what kind of adventure he\'ll find there! When DK, Cranky Kong, and Diddy Kong reached the top, they encountered Xananab, an alien that looked like a banana. But they also saw King K. Rool and his four Kremling advisors making off with the five Crystal Bananas, five objects Xananab wanted back. DK agreed to help Xananab get the Crystal Banana back, thus starting off his next adventure, with Diddy at his side.
Donkey and Diddy (along with Cranky and Xananab) travel through several islands, including Ghost Island, Lost Island, and Chill \'n\' Char Island. At the end of the last level of each island DK had to fight a Kremling mutated by one of the Crystal Bananas, and going into a machine. After beating the boss, they gained a Crystal Banana. At the end of Chill \'n\' Char Island, after the boss, K. Rool and his final Kremling make way to the King Kruizer IV, an updated model of K. Rool\'s cruiser seen in Donkey Kong Country 2/Donkey Kong Land 2, Donkey Kong 64, and DK: King of Swing. They travel to the top of a very large island, High-High Island, to make it just a little too late. Fortunately, Cranky gives them a Booster Barrel (also seen in King of Swing), which they use to travel into the King Kruizer IV. Once inside, they travel through the vehicle, and DK has to fight the final Kremling, gaining a Crystal Banana once victorious.
K. Rool flees with his last device, going into a wormhole, which eventually leads DK and friends into Xananab\'s homeplanet, Planet Plataen. K. Rool is fought here, and, once defeated, uses the final Crystal Banana to mutate and become gigantic. Once DK defeats K. Rool in this state, K. Rool is defeated and the game ends. Xananab thanks DK, Diddy, and Cranky by making them the local celebrities and they are able to eat all the bananas they can eat. Eventually, the three monkeys head home, and humorously tie K. Rool to the back of the Banana Spaceship. It is not revealed what happens when they return home.